The sperm nucleus observed in the embryo sac of Tricyrtis hirta (Liliaceae)

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ユリ科植物のホトトギスを材料として、花粉管が到達する前の胚嚢と到達後の胚嚢を観察した。花粉管が到達する前に、中央細胞内の2個の極核は融合し、大きな1個の複相の核になる。また、卵核と複相核は球形をしており、核膜は張りきっている。しかし、この両核は時間の経過と共に次第に変化し、花粉管が助細胞に到達し、精核が卵細胞と中央細胞に入ったときには、卵核と複相核の核膜は波を打ったようにたるみが出来る。卵細胞と中央細胞に入った直後の精核は楕円体をしていると思われる。精核が卵核や複相核に近づくにつれて、卵核や複相核に面している精核の側面が湾曲の程度を減少させ、精核の形がいびつになる。精核の側面の変形に合わせて、精核に面している卵核と複相核の核膜は精核の変形した核膜に並行するように変わる。精核が卵核と中央細胞の複相核に接着する直前まで、精核と卵核、複相核のそれぞれの核膜は2重膜構造を維持している。
The embryo sac before a pollen tube reaches, and the embryo sac after a pollen tube reaches were observed in Tricyrtis hirta of liliaceaous plant. Before the pollen tube reaches the embryo sac, two polar nuclei inside the central cell fuse, and become one large diploid nucleus. Moreover, an egg nucleus and the diploid nucleus have a globular form, and a nulear envelope of them spreads completely. But, looseness like a wave occurs in the nulear envelope of the egg nucleus and the diploid nucleus, when the pollen tube reached the embryo sac and a sperm nucleus was included in the egg cell and the central cell. The sperm nucleus immediately after it went into the egg cell and the central cell is thought to have an ellipsoid. As the sperm nucleus approaches the egg nucleus and the diploid nucleus, the side of the sperm nucleus which faces the egg nucleus and the diploid nucleus decreases in the degree of curving, and the shape of the sperm nucleus warps. The nulear envelope of the egg nucleus to face the sperm nucleus changes to go side by side with the nulear envelope which warped in the sperm nucleus in accordance with the deformation on the side of the sperm nucleus. The nulear envelope of the sperm nucleus, the egg nucleus and the diploid nucleus of the central cell maintains a double membrane structure while the sperm nucleus is not contact with to the egg nucleus and the diploid nucleus of the central cell.

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